Infection of A549 cells with a recombinant LCMV-Armstrong engineered to express GFP, LCMV-2A-GFP, also show a marked reduction in infection upon inactivation of CD164, ST3GAL4, or SLC35A1 (Fig. 2B and SI Appendix, Fig. S2D). This evidence concerns the gene ST3GAL4 and infection.