Genes SLC22A4, OTUD3, PARK7, NOTCH4, GPR35, DAP, UBA7, MST1, MST1R, CD40, TAP2, NICNI, RGS14, LINCO1475, GBAP1, NUCKS1, HCG23, CCDC88B, HEART3, and ERAP2, have been previously associated with IBD, CD or UC. Here, CCDC88B is linked to inflammatory bowel disease.