The remaining variants rs56402156, rs7920721, and rs4351014 (in/near EPHA1, ECHDC3, and HS3ST1) have not been directly associated with other traits, although their associated genes were implicated in systemic lupus erythematosus (HS3ST1) and cancer (EPHA1, ECHDC3) (Herath et al., 2009; Rafiq et al., 2014; Langefeld et al., 2017). Here, EPHA1 is linked to systemic lupus erythematosus.