HLA-DQB1 normalized read counts from uninflamed colon biopsies (n = 28) from healthy control subjects or patients with Crohn’s disease or ulcerative colitis were stratified according to DQA1*01:02 carrier status and genotype at the SNP rs3135006, which is in LD with rs17612852 in the LEAP cohort (D′ = 0.98, r2 = 0.79). Here, HLA-DQA1 is linked to Crohn disease.