According to the predictive database miRWalk [119], all four differentiation markers are predicted to be targeted by IBD-associated miR-16, miR-106, miR-21 (excluding KLF4 and HATH1), miR-122 (excluding KLF4), miR-146, miR-151, miR-155 (excluding KLF4, GFI1 and SPDEF), miR-199 (excluding GFI1) and miR-362 (excluding GFI1 and HATH1). Here, GFI1 is linked to inflammatory bowel disease.