MTO1 and hepatocellular carcinoma: The pull-down assay results showed that the expressions of circMTO1, miR-541-5p, and miR-9 were enriched in HCC cell lines compared to that in controls, and the expression of miR-541-5p was relatively greater than that of miR-9, whereas the linear MTO1 had no enrichment (Fig. 2C, D).