The function is fulfilled by an oncogenic complex composed of BRD3, NSD3, and ZNF532 (and other zinc finger proteins, ZNF592, ZNF687, and ZMYND8, collectively termed ‘Z4’ [79]), which have all been identified as NUT fusion partners in a subset of NUT carcinoma [80]. This evidence concerns the gene NUTM1 and nut midline carcinoma.