Furthermore, normal cells have modifications in long-branched sugar chains, whereas cancer cells express various simple and short sugar chain antigens called O-glycans (e.g., Tn, sialyl-Tn and sialyl-Lewis-X), these antigens were reported to have strong binding signals with Gg and Rn, making it an interesting avenue for future applications of Gg and Rn to study cancer cells. The gene discussed is FUT4; the disease is cancer.