MUC5AC and cancer: In addition, unusual expression of mucin-type O-glycans has been associated with several human disorders, including Tn syndrome (Berger, 1999; Ju and Cummings, 2005), IgA nephropathy (Suzuki et al., 2008; Hiki, 2009), heterotaxy (Fakhro et al., 2011; Boskovski et al., 2013), and cancer (Springer, 1984; Ju et al., 2008; Radhakrishnan et al., 2014).