TUBB4B and neoplasm: Furthermore, we quantified a clear enrichment of cilia genes such as ran, tubb4b, tuba4l, and gmnn specifically in tumor-like and muscle-like interface cells in our snRNA-seq dataset, and, similar to our scRNA-seq results, all four genes were upregulated more highly in the muscle-like interface cluster than in any of the other interface clusters (Fig. 5f).