Finally, SNPs that were molecular QTLs in our study colocalized with asthma GWAS SNPs, revealing 46 unique colocalizations that included both known asthma loci (e.g., 17q12-21 and TSLP) and loci that did not meet stringent criteria for genome-wide significance in the GWASs (e.g., IRF5 and PMM1) (Additional file 10: Table S4). Here, IRF5 is linked to asthma.