SGCG and lung cancer: Jabalameli et al. studied the carriage of Type III secretory toxin-encoding genes and their cytotoxic action on A549 human lung cancer cell lines in 96 clinical P. aeruginosa isolates, finding that the carriage of exoT, exoY, exoU, and exoS were 100%, 95%, 64.5%, and 29%, respectively.