We identified FDR-significant (p < ~5 × 10−3) enrichment for all four diseases in CD4+ T cells in lung, and for MS, IBD and CD in CD8+ cytotoxic T cells in both lung and spleen, and regulatory T cells in lung (Fig. 5, Figs. S16–18, S21–23, S26–28, S31–33, Supplementary Data 4). Here, CD4 is linked to inflammatory bowel disease.