The global and gene-specific (DICER1, FGF2, TACC1, RAB10, RUNX121, ZEB1, PDXK, COL1A2, PPP3CB, C6orf89, SAP30L, APH1B, CHURC131) shortening observed in our study is consistent with previous findings in other cancer types using different approaches21,31,46. Here, FGF2 is linked to cancer.