Open-ended LC-MS analysis indicated that both mouse and human H19 sense transcripts, but not anti-sense transcripts, associated with DMD complex components, including DMD, NOS1 (Nitric oxide synthase), SNTA1 (syntrophin alpha 1), DAG1 (beta-dystroglycan), SGCA (alpha-sarcoglycan), and SNTB1/2 (Beta-1-syntrophin/ Beta-2-syntrophin) [28] (Fig. 1a and Additional file 5: Table S4). Here, DAG1 is linked to Duchenne muscular dystrophy.