In addition, “cilium morphogenesis” emerged as an enriched term (DAVID enrichment score 0.87) and several genes associated with RA risk SNPs in FLS were connected to the formation of the primary cilium (C5orf30, GSN, TMEM138, TMEM216, CNTRL, INCENP, ACTR2). Here, TMEM216 is linked to rheumatoid arthritis.