MAPT and supranuclear palsy, progressive, 1: Furthermore, this study revealed that tau filaments in an atypical PSP case with limbic globular 4R-positive neuronal inclusions had a unique structure intermediate between GGT and PSP tau filaments, providing molecular evidence to define this case as a novel class of tauopathy, which they termed as Limbic-predominant Neuronal inclusion body 4R Tauopathy (LNT) [167].