Cep290 (120, 121) and Inpp5e (19), associated with syndromic ciliopathies that include obesity, and Bardet-Biedl syndrome (BBS) causal genes, Arl6, Bbs1, Bbs2, Bbs5, Bbs7 (122), were also present in these modules. Here, CEP290 is linked to ciliopathy.