In addition to the arachidonic acid QTL that colocalized with Crohn’s disease, the FUT2 locus led by non-synonymous variant rs601338 also showed colocalization with three immune-mediated diseases: Celiac disease [59] (Coloc analysis H4: 0.7, H3: 0.004), Crohn’s disease [44] (Coloc analysis H4: 0.98), and type 1 diabetes (Coloc analysis H4: 0.99) [60]. Here, FUT2 is linked to celiac disease.