ST3GAL4 and membranous glomerulonephritis: In summary, we identified 13 genome-wide significant suPAR-associated variants and based on literature searches and GO term annotations, the variants are found in and around 12 genes encoding uPAR/suPAR (PLAUR) and its ligand uPA (PLAU), genes with relations to glycoprotein biosynthesis and glycosylation (ASGR1, ST3GAL4, ST3GAL6, ABO, CHST9, FUT2), genes involved in immune response (LY75, HLA-DRA, CFH), and PLA2R1 (of which variants have been previously associated with membranous nephropathy).