Among the various factors in cancers, transforming growth factor (TGF)-β is a major cytokine that induces invasion and metastasis through an EMT (Behrens et al., 1989; Buck and Knabbe, 2006; Barcellos-Hoff and Akhurst, 2009; Drabsch and ten Dijke, 2012). Here, TGFB1 is linked to cancer.