RTL9 and metabolic syndrome: The inclusion of the X chromosome within the study allowed us to identify an intronic variant (rs67648651) associated with dyslipidemia in the RGAG1 gene among the previously known associated loci since a variant in LD with our top variant (rs5985471, LD r2 = 0.92 for European ancestry) has already been associated with low-density lipoprotein cholesterol20.