While FOXA1 and GATA3 are known to have low expression in basal bladder cancer cell lines and tumors, the enrichment of forkhead and GATA motifs in open chromatins across BLCA cell lines suggests compensation by Forkhead and GATA factors other than FOXA1 and GATA3. Here, QRSL1 is linked to bladder transitional cell carcinoma.