They identified co-localization of eQTL top variant and SLE GWAS variant in 9 genes (UBE2L3, HLA-DRB5, RP11-356I2.3, BLK, FAM167A, NADSYN1, RP11-660L16.2, ALDH2, and ALDH18A1), and 13 of the 26 genes with perturbations in both expression and splicing in SLE had a splicing QTL. This evidence concerns the gene UBE2L3 and systemic lupus erythematosus.