In these legumes, the infection is orchestrated by several transcription factors, including nodule inception (Nin; Schauser et al., 1999; Marsh et al., 2007), Nsp1/Nsp2 (Kalo et al., 2005; Heckmann et al., 2006), Cyclops (Yano et al., 2008; Singh et al., 2014), and ERF required for nodulation (Ern1; (Cerri et al., 2012; Kawaharada et al., 2017). Here, SH2D3A is linked to infection.