In contrast, although cancer cell expression of the antisense noncoding RNA in the INK4 locus (ANRIL) and of the lymph node metastasis associated transcript 1 (LNMAT1) have been associated with lymphangiogenesis and lymphatic metastasis29,30, lymphatic endothelial-specific lncRNAs have not been identified or functionally characterized so far. Here, DUXAP9 is linked to cancer.