For breast cancer GWAS35, enriched in epithelial and cancer biosamples (Extended Data Fig. 9c), the highly localized rs17356907 genetic signal (P = 10−39, rank no. 12) localized precisely in a narrow epithelial and cancer enhancer nearest to USP44 but linked instead to NTN4, which is implicated in tumorigenesis and angiogenesis (Extended Data Fig. 9d). Here, NTN4 is linked to cancer.