TRH and hemorrhagic disease: Seven Vibrio strains (including V. vulnificus HM-TA-D2-L2-V2; V. vulnificus HM-TA-G2-V1-D2; V. brasiliensis HM-X-13/6; V. cholerae V-13/6; V. parahaemolyticus HM-17/6; V. cholerae HM-V-13/6 and V. vulnificus HM-X-13/6) causing hemorrhagic disease in red drum (S. ocellatus) had only the tlh gene, and none of the Vibrio strains had tdh and trh genes (Quang et al. 2020).