We found that a number of TFs were encompassed by BEC-SEs including TP63, a well-defined marker of basal cells, and TP73, a key player in mucociliary development (Supplementary Figure S1E) and observed enrichment for FOXI1 sites in asthma DERs, a TF shown to characterize the recently identified ionocytes (Montoro et al., 2018; Plasschaert et al., 2018). Here, TP63 is linked to asthma.