In addition, variants in FERMT (rs17125944), NCAPD2 (rs740850), and TESC (rs7294919) were associated with brain amyloidosis (Apostolova et al., 2018), AD (Lee et al., 2008), and hippocampal volume/brain development (Stein et al., 2012). Here, NCAPD2 is linked to Alzheimer disease.