Conversely, markers for PNs (Emx2), oligodendrocytes (Olig1 and Olig2) and astrocytes (Aldh1l1) showed greater enrichment of H3K27ac over the gene body and proximal sequences in the non-CIN samples (Fig. 1C–E). Here, EMX2 is linked to cervical squamous intraepithelial neoplasia.