Six significant SNPs were associated with MCHC, three of these were located within genes including VPS13D (vacuolar protein sorting 13 homolog D) on Oar12, OTOA (otoancorin) on Oar24, and RSU1 (Ras suppressor protein 1) on Oar13, and the others were located from 67,432 to 496,548 bp from the nearest gene. This evidence concerns the gene RSU1 and mean corpuscular hemoglobin concentration.