HLA-DRB1 and leprosy: For this, we adjusted each of the formally non-significant HLA alleles on the presence of HLA-DQA1*01:05, HLA-C*07:06 and HLA-DRB1*12 and identified seven HLA four-digit alleles with evidence (Pconditional < 0.01) for association with leprosy (S5 Table).