In PD and DLB, α-synuclein pathologies are mainly observed in neurons in the form of Lewy bodies (LBs) and Lewy neurites (LNs) (Baba et al., 1998), while glial cytoplasmic inclusions (GCIs) are seen in oligodendrocytes in MSA (Wakabayashi et al., 1998). Here, SNCA is linked to multiple system atrophy.