While the significance of TERT—BRG1 association in Wnt signaling remains to be further investigated, the heterotrimeric complex consisting of TERT, BRG1, and the nucleolar protein nucleostemin—a GNL3L (Section 3.1) paralog that likely does not participate in TERT catalytic action at telomeres [69]—has been studied extensively, with implicated non-telomeric functions including tumor initiation, heterochromatin assembly, and microRNA regulation [107,108,109]. Here, SMARCA4 is linked to neoplasm.