S100A4 and cancer: We also detected enrichment for transcripts encoding fibroblast-specific protein 1 (FSP1/S100a4), keratins Krt7, Krt8, Krt14 and Krt18 and claudins Cldn3, Cldn4 and Cldn7 in this cluster which were also enriched in epithelial/cancer cells (Figure 3F and Figure S2B,D).