KIF4A and microcephaly: Furthermore, in addition to Citron kinase and RhoA, three members of Kinesin family, Kif20a, Kif20b, and Kif4 regulate neocortical NPC cytokinesis and mutations of these genes result in reduced cortical size with microcephaly (Janisch et al., 2013; Moawia et al., 2017; Geng et al., 2018).