Seventeen variants were reported in genes that had been previously associated with hypospadias (ATF3, BNC2, CYP1A1, EMX2, EYA1, FLNA, GLI3, GRID1, GLI2, BNC2, FGF10, HOXA13, HSD3B2, IRX5, IRX6, PPARGC1B, and WDR11), eight with cryptorchidism (BNC2, FLNA, RET, RECQL4, NRP1, PTPN11, RIPK4, and ZBTB16), and five with micropenis (ZBTB16, BNC2, EVC, FGF10, and RIPK4). Here, RIPK4 is linked to cryptorchidism.