A total of eight genes detected in GWAS signals were also successfully fine-mapped, including Plexin A4 (PLXNA4), FA Complementation Group C (FANCC), Neurotrimin (NTM) for displaced abomasum, GC for mastitis and livability, ATP Binding Cassette Subfamily C Member 9 (ABCC9) for livability, QDPR for retained placenta, Zinc Finger And AT-Hook Domain Containing (ZFAT) and CCDC88C for livability. The gene discussed is ABCC9; the disease is mastitis.