FOXO4 and vitamin A deficiency: Our team has shown that embryonic expression of the lncRNA SULT1C2A was decreased whereas miR‐466c‐5p was upregulated in the rat model of vitamin A deficiency‐induced congenital scoliosis.43 Somitogenesis‐related genes, such as Foxo4, Sox9, Pax1 and Nkx3‐2, were downregulated on gestational day 9 upon maternal vitamin A deficiency.