In addition, we identified recurrent mutations in two genes involved in actomyosin dynamics in ASL and HCC: COL1A1 (Collagen Type I Alpha 1 Chain) and GMIP (GEM Interacting Protein), both of which have been previously reported as mutated in HCC29–31, but have not been previously described in ASL. Here, GMIP is linked to hepatocellular carcinoma.