SFRP4 and benign prostatic hyperplasia: Twelve PYLs carrying two BPH resistance genes (BPH2 + BPH17-PYL, BPH2 + BPH25-PYL, BPH2 + BPH32-PYL, BPH2 + BPH17-ptb-PYL, BPH3 + BPH17-PYL, BPH17 + BPH21-PYL, BPH20 + BPH21-PYL, BPH20 + BPH32-PYL, BPH21 + BPH25-PYL, BPH21 + BPH17-ptb-PYL, BPH25 + BPH17-ptb-PYL and BPH32 + BPH17-ptb-PYL) and three PYLs containing three BPH resistance genes (BPH2 + BPH3 + BPH17-PYL, BPH2 + BPH32 + BPH17-ptb-PYL and BPH20 + BPH21 + BPH32-PYL) were developed using NILs and PYLs with BPH resistance gene(s) (Table 2).