TARDBP and amyotrophic lateral sclerosis: Finally, the genes analyzed for ALS were C9orf72 (C9orf72-SMCR8 complex subunit); TARDBP (TAR DNA Binding Protein), SOD1 (Superoxide Dismutase 1); FUS (FUS RNA Binding Protein), UBQLN2 (Ubiquilin 2) and HNRNPA2B1 (Heterogeneous Nuclear Ribonucleoprotein A2/B1).