These include the CpG in LMAN2 (cg23483530) in the sienna3 module, which presents the second strongest negative association with the MSA status, and the CpG in HIP1 (cg15769835) in the lightcyan module, which was positively associated with the MSA OPCA subtype. Here, LMAN2 is linked to olivopontocerebellar atrophy.