Both interannual mean density (Fig 2A) and biomass (± standard error; Fig 2C) were highest in CH (76.80 ± 8.40 no./100m2; 266.18 ± 11.46 g/100m2), followed by TB (40.43 ± 4.42 no./100m2; 103.18 ± 4.79 g/100m2), AB (23.17 ± 2.39 no./100m2; 89.19 ± 5.48 g/100m2), then CK (6.37 ± 0.93 no./100m2; 38.64 ± 3.45 g/100m2). This evidence concerns the gene C4B and tuberculosis.