In addition, the inter-clade recombination included sequences resembling both syphilis and bejel sequences (in TP0117 coding for TprC; in TP0317 coding for TprG; in TP0462 encoding probable lipoprotein; in TP0483 coding for hypothetical protein; in TP0621 coding for TprJ; in TP0865 encoding a putative outer membrane protein) (Table 2 and Figure 3). Here, PRCC is linked to syphilis.