ADIPOQ and metabolic syndrome: In contrast, the stable enhancers are strongly associated with liver-related traits such as metabolic syndrome (blood metabolic levels, blood metabolic ratios), blood lipid levels (LDL peak particle diameter, adiponectin levels, visceral adipose tissue adjusted for BMI, visceral adipose tissue, subcutaneous adipose tissue ratio), and glucose homeostasis traits (Additional file 2: Tables S1-S4 and Fig. 3b).