A less prominent QTL on SSC4 harbors variants in or close to the aforementioned genes RPS20 (17 downstream variants, min p-value 1.51 × 10−25) and in SNTG1 (19 intron variants, min p-value 1.48 × 10−14), which has been associated with type 2 diabetes (Ban et al. 2010). The gene discussed is RPS20; the disease is type 2 diabetes mellitus.