To generate the Hipk tumor model, we used the Gal4-UAS system (Brand and Perrimon, 1993) to induce overexpression of hipk in larval wing discs (full genotype: dpp-Gal4>UAS-RFP + UAS-hipk, abbreviated as dpp>RFP + hipk) (Figure 1c–d). This evidence concerns the gene LGALS4 and neoplasm.