Thirteen CpG were positively associated with hypercholesterolemia: cg08862778 (MTOR), cg07199894 (ULK1), cg11658986 (ADCY6), cg04149773 (ADCY6), cg06223834 (ADCY9), cg18237616 (RHEB), cg05792022 (FOXO1), cg01781374 (CAMK4), cg11301281 (CREB5), cg14267811 (TSC1), cg08128650 (RELA), cg01284192 (IGF1R), cg24061580 (PRKAG2) (Figure 2). Here, FOXO1 is linked to familial hypercholesterolemia.